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From Biology-Online Dictionary
In a subtle manner; slyly; artfully; cunningly. Thou seest how subtly to detain thee I devise. (milton)
2. Nicely; delicately. In the nice bee what sense so subtly true. (pope) Subtly communicating itself to my sensibilities, but evading the analysis of my mind. (Hawthorne)
3. Deceitfully; delusively.