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From Biology-Online Dictionary
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1. (Science: mathematics) More, required to be added; positive, as distinguished from negative; opposed to minus.

2. Hence, in a literary sense, additional; real; actual. Success goes invariably with a certain plus or positive power. (Emerson)

3. (Science: mathematics) Plus sign, the sign (_) which denotes addition, or a positive quantity.

Origin: L, more; akin to Gr, and cf. Piu, Pleonasm.