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From Biology-Online Dictionary
1. Blame; cause of blame; fault; crime; offense.
2. Deficiency; want; need; destitution; failure; as, a lack of sufficient food. She swooneth now and now for lakke of blood. (Chaucer) Let his lack of years be no impediment. (Shak)
Origin: oe. Lak; cf. D. Lak slander, laken to blame, OHG. Lahan, as. Lean.