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From Biology-Online Dictionary
The tincture red, indicated in seals and engraved figures of escutcheons by parallel vertical lines. Hence, used poetically for a red colour or that which is red. His sev'n-fold targe a field of gules did stain in which two swords he bore; his word, Divide and reign.' (P. Fletcher) Follow thy drum; With man's blood paint the ground; gules, gules. (Shak) Let's march to rest and set in gules, like suns. (Beau. & Fl)
Origin: oe. Goules, f. Gueules, the same word as gueule throat, OF. Gole, goule, L. Gula. So named from the red colour of the throat. See gullet, and cf. Gula.