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From Biology-Online Dictionary
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1. Next in order after the fourth; the ordinal of five.

2. Consisting of one of five equal divisions of a thing. Fifth monarchy men, a fanatical sect in England, of the time of the commonwealth, who maintained that there would be a fifth universal monarchy, during which christ would reign on earth a thousand years. Fifth wheel, a horizontal wheel or segment above the fore axle of a carriage and beneath the body, forming an extended support to prevent careening.

Origin: oe. Fifte, fifthe, as. Fifta. See five.