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From Biology-Online Dictionary | Biology-Online Dictionary
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1. To breathe out. Hence: to emit, as vapor; to send out, as an odour; to evaporate; as, the earth exhales vapor; marshes exhale noxious effluvia. Less fragrant scents the unfolding rose exhales. (Pope)

2. To draw out; to cause to be emitted in vapor; as, the sum exhales the moisture of the earth.

Origin: L. Exhalare; ex out _ halare to breathe; cf.F. Exhaler. Cf. Inhale.