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<p>The Aubesier 4, 10, and 11 human remains therefore provide further support for the gradual increase in frequency of derived Neandertal features in the northwestern Old World during the Middle Pleistocene, most likely as a product of isolation by distance from contemporaneous Africa and eastern Asian human populations. At the same time, these remains reinforce several paleobiological aspects of these human populations, including their frequent nondietary use of the anterior dentition and the prolonged survival of individuals with serious impairments. These human populations therefore had achieved a level of sociocultural elaboration sufficient to maintain debilitated individuals and to provide the motivation to do so. However, they also were using their biology to accomplish daily activities, reflecting behavioral patterns and a level of technological development significantly different from those of Upper Paleolithic and recent humans. </p>
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