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ACKNOWLEDGEMENTS<br><br> This work was funded through two grants from NIAID/DARPA: U01 AI48706 (E.J.L.) and U01 AI48653-02 (R.M.L.B. and C.U.) and Canadian NSERC grant OPG0155125-01 to C.U. <p> <br> Notes<br><br> The online version of this article has been published under an open access model. Users are entitled to use, reproduce, disseminate, or display the open access version of this article for non-commercial purposes provided that: the original authorship is properly and fully attributed; the Journal and Oxford University Press are attributed as the original place of publication with the correct citation details given; if an article is subsequently reproduced or disseminated not in its entirety but only in part or as a derivative work this must be clearly indicated. For commercial re-use permissions, please contact journals.permissions@oupjournals.org. </p>
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