Ecologists have long been intrigued by the factors that control the pattern …

• Abstract
• Results
• Discussion
• Materials and Methods
• Acknowledgements
• References
• Figures
• Tables

  
Hubbell, SP. (2001) The Unified Neutral Theory of Biodiversity and Biogeography (Princeton Univ Press, Princeton,). Preston, FW. (1948) Ecology 29, 254–283 Lewinsohn, TM, Novotny, V & Basset, Y. (2005) Annu Rev Ecol System 36, 597–620 Andow, DA. (1991) Annu Rev Entomol 36, 561–586 Murdoch, MW, Evans, FC & Peterson, CH. (1972) Ecology 53, 819–829 Novotny, V, Drozd, P, Miller, SE, Kulfan, M, Janda, M, Basset, Y & Weiblen, GD. (2006) Science 313, 1115–1118 Armbrecht, I, Perfecto, I & Vandermeer, J. (2004) Science 304, 284–286 Haddad, NM, Tilman, D, Haarstad, J, Ritchie, M & Knops, JMH. (2001) Am Nat 158, 17–35 Siemann, E, Tilman, D, Haarstad, J & Ritchie, M. (1998) Am Nat 152, 738–750 Hunter, MD. (1992) in Effects of Resource Distribution on Animal–Plant Interactions eds. Humter, MD, Ohgushi, T& Price, PW. (Academic, New York,) pp. 287–325. Kelly, CK & Southwood, TRE. (1999) Proc Natl Acad Sci USA 96, 8013–8016 Marques, ESDA, Price, PW & Cobb, NS. (2000) Environ Entomol 29, 696–703 Ohgushi, T. (1992) in Effects of Resource Distribution on Animal–Plant Interactions eds. Hunter, D, Ohgushi, T& Price, PW. (Academic, New York,) pp. 287–325. Potts, SG, Vulliamy, B, Dafni, A, Neåfeman, G & Willmer, P. (2003) Ecology 84, 2628–2642 Schoener, TW. (1974) Science 185, 27–39 Tokeshi, M. (1999) Species Coexistence: Ecological and Evolutionary Perspectives (Blackwell, Oxford,). Gilbert, B & Lechowicz, MJ. (2004) Proc Natl Acad Sci USA 101, 7651–7656 Bell, G. (2005) Ecology 86, 1757–1770 Dornelas, M, Connolly, SR & Hughes, TP. (2006) Nature 440, 80–82 Graves, GR & Rahbek, C. (2005) Proc Natl Acad Sci USA 102, 7871–7876 McGill, BJ. (2003) Nature 422, 881–885. Tuomisto, H, Ruokolainen, K & Yli-Halla, M. (2003) Science 299, 241–244 Manly, BFJ. (1991) Randomization and Monte Carlo Methods in Biology (Chapman & Hall, London,). Manly, BFJ. (1986) Res Pop Ecol 28, 201–218 Soininen, J, McDonald, R & Hillebrand, H. (2007) Ecography 30, 3–12. Kitahara, M & Watanabe, M. (2003) Ecological Research 18, 503–522 Usher, MB & Keiller, SW. (1998) Biodiversity Conservation 7, 725–748 Steffan-Dewenter, I & Tscharntke, T. (1997) Oecologia 109, 294–302 Hooper, DU, Chapin, I, Ewel, JJ, Hector, A, Inchausti, P, Lavorel, S, Lawton, JH, Lodge, DM, Loreau, M & Naeem, S, et al. (2005) Ecol Monogr 75, 3–35 Fukuda, H, Haba, E, Kuzuya, K, Takahasi, A, Takahasi, M, Tanaka, B, Tanaka, H, Wakabayashi, M & Watanabe, Y. (1982) The Life Histories of Butterflies in Japan (Hoikusha, Osaka, Japan,) Vol 1–4 (in Japanese). Quinn, GP & Keough, MJ. (2002) Experimental Design and Data Analysis for Biologists (Cambridge Univ Press, Cambridge,). Legendre, P & Legendre, L. (1998) Numerical Ecology (Elsevier Scientific Pubishing, Amsterdam,). Benjamini, Y & Hochberg, Y. (1995) J R Stat Soc Ser B 57, 289–300.