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Fossil land mammals from ten localities of southern Buenos Aires Province, Argentina …

Biology Articles » Paleobiology » Late Cenozoic mammal bio-chronostratigraphy in southwestern Buenos Aires Province, Argentina » Appendix

- Late Cenozoic mammal bio-chronostratigraphy in southwestern Buenos Aires Province, Argentina

Appendix 1. Systematic list of the taxa found in the study area, up to the levels in which the new materials could be determined / Lista sistemática de los taxones hallados en el área de estudio, hasta el nivel en que los nuevos materiales pudieron ser determinados.

Class Osteichthyes Howes, 1894
Order Cyprinodontiformes Bertin, 1958
Cyprinodontiformes indet.
Order Siluriformes Fowler, 1951
Family Pimelodidae Eigenmann and Eigenmann, 1889
Genus Pimelodella Eigenmann, 1917
P . aff. laticeps (in Cione and López Arbarello, 1995)
Family Callichthyidae Gill, 1872
Genus Callichthys Linné, 1758
C. callichthys (in Cione and López Arbarello, 1995)
Genus Corydoras Lacépède, 1803
Corydoras cf. C. paleatus (Jenyns, 1842)
Order Perciformes Ludwig, 1883
Family Percichthyidae Jordan, 1923
Genus Percichthys Girard, 1854

Class Amphibia Linné, 1758
Order Anura Giebel, 1847
Anura indet.

Class Reptilia Laurenti, 1768
Order Chelonii Brongniart, 1800
Family Chelidae Gray, 1825
Genus Hydromedusa Wagler, 1830
H. tectifera (Cope, 1869)
Order Squamata Oppel, 1811
Family Iguanidae Gray, 1827
Iguanidae indet.
Class Aves Linné, 1758
Order Rheiformes Fürbringer, 1888
Family Rheidae Bonaparte, 1853
Genus Rhea Brisson, 1760
R. americana (Linné, 1758)
Order Tinaniformes (Huxley, 1872)
Family Tinamidae Gray, 1840
Genus Nothura Wagler, 1827
N . darwini Gray, 1867
Order Anseriformes Garrod, 1874
Family Anatidae Vigors, 1825
Genus Anas Linné, 1758
A. platalea Vieillot, 1816
Genus Chloephaga Eyton, 1838
Chloephaga sp. 1
Chloephaga sp. 2
Genus Dendrocygna Swinson, 1837
Order Gruiformes (Bonaparte, 1854)
Family Rallidae Vigors, 1825
Genus Porphyrula Linné, 1758
Order Passeriformes (Linné, 1758)
Suborder Tyranni Wetmore and Miller, 1926
Family Furnariidae (Gray, 1840)
Subfamily Philydorinae Sclater, 1890
Genus Pseudoseisura Reichenbach, 1853
Genus Pseudoseisuropsis Noriega, 1991
Suborder Passeres Linné, 1766
Family Passeridae (Illiger, 1811)
Subfamily Motacillinae Bonaparte, 1831

Class Mammalia Linné, 1758
Superorder MarsupialiA Illiger, 1811
Order Sparassodonta (Ameghino, 1894)
Family Hathliacynidae Ameghino, 1894
Genus Borhyaenidium Pascual and Bocchino, 1963
Order Didelphimorphia Gill, 1872
Family Didelphidae Gray, 1821
Subfamily Marmosinae Reig, 1981
Genus Lestodelphys Tate, 1934
Lestodelphys halli (Thomas, 1921)
Genus Thylamys Gray, 1843
T. pusillus (Desmarest, 1804)
Superorder Xenarthra Cope, 1889
Order Cingulata Illiger, 1811
Superfamily Dasypodoidea Gray, 1821
Family Dasypodidae Bonaparte, 1838
Subfamily Dasypodinae Bonaparte, 1838
Tribe Dasypodini Bonaparte, 1838
Genus Propraopus Ameghino, 1881
Subfamily Euphractinae Pocock, 1924
Tribe Eutatini Bordas, 1933
Genus Doellotatus Bordas, 1932
Doellotatus inornatus (Rovereto, 1914)
Doellotatus praecursor (Rovereto, 1914)
Genus Chasicotatus Scillato Yané, 1979
Chasicotatus peiranoi Esteban and Nasif, 1996
Genus Eutatus Gervais, 1867
Eutatus seguini Gervais, 1867
Tribe Euphractini Pocock, 1924
Genus Chorobates Reig, 1959
Chorobates villosissimus (Rovereto, 1914)
Genus Macrochorobates Scillato Yané, 1980
Genus Macroeuphractus Ameghino, 1887
Macroeuphractus morenoi (Lydekker, 1894)
Genus Chaetophractus Fitzinger, 1871
Chaetophractus villosus (Desmarest, 1804)
Genus Zaedyus Ameghino, 1889
Zaedyus pichiy (Desmarest, 1804)
Subfamily Tolypeutinae Gray, 1865
Tribe Tolypeutini Gray, 1865
Genus Tolypeutes Illiger, 1811
Tolypeutes matacus (Desmarest, 1804)
Superfamily Glyptodontoidea Gray, 1869
Family Glyptodontidae Burmeister, 1879
Subfamily Sclerocalyptinae Ameghino, 1895
Tribe Sclerocalyptini Ameghino, 1895
Genus Sclerocalyptus Ameghino, 1891
Sclerocalyptus ornatus (Owen, 1845)
Genus Berthawyleria Castellanos, 1939
Tribe Palaehoplophorini Hosffstetter, in Piveteau, 1958
Genus Aspidocalyptus Cabrera, 1939
Tribe Lomaphorini Hoffstetter, 1958
Tribe Plohophorini Castellanos, 1932
Genus Plohophorus Ameghino, 1887
Plohophorus cuneiformis Ameghino, 1904
Tribe Panochthini Castellanos, 1927
Genus Panochthus Burmeister, 1866
Panochthus tuberculatus (Owen, 1839)
Tribe Neuryurini Hoffstetter, 1958
Subfamily Doedicurinae Ameghino, 1889
Genus Doedicurus Burmeister, 1874
Subfamily Glyptodontinae Gray, 1869
Tribe Glyptodontini Gray, 1869
Genus Glyptodon Owen, 1839
Glyptodon clavipes Owen, 1839
Order Tardigrada Lathan y Davies, 1795
Family Mylodontidae Gill, 1872
Subfamily Scelidotheriinae Ameghino, 1889
Genus Scelidotherium Owen, 1839
Scelidotherium leptocephalum Owen, 1839
Subfamily Mylodontinae Gill, 1872
Genus Glossotherium Owen, 1839
Genus Lestodon Gervais, 1855
Lestodon armatus Gervais, 1855
Family Megatheriidae Owen, 1843
Genus Megatherium Cuvier, 1796
Megatherium americanum Cuvier, 1796
Order Litopterna Ameghino, 1889
Family Proterotheriidae Ameghino, 1887
Subfamily Proterotheriinae Ameghino, 1885
Genus Epitherium Ameghino, 1888
Epitherium laternarium Ameghino, 1888
Family Macraucheniidae Gervais, 1855
Genus Promacrauchenia Ameghino, 1904
Genus Macrauchenia Owen, 1838
Macrauchenia patachonica Owen, 1838
Genus Macraucheniopsis Paula Couto, 1945
Macraucheniopsis ensenadensis (Ameghino, 1888)
Order Notoungulata Roth, 1903
Family Toxodontidae Owen, 1845
Genus Toxodon Owen, 1837
Family Mesotheriidae Alston, 1876
Subfamily Mesotheriinae Alston, 1876
Genus Pseudotypotherium Ameghino, 1904
Family Hegetotheriidae Ameghino, 1894
Subfamily Pachyrukhinae Kraglievich, 1934
Genus Paedotherium Burmeister (in C.V. Burmeister, 1888)
P. minor Cabrera, 1937
P. bonaerense (Ameghino, 1887)
Genus Tremacyllus Ameghino, 1891
T. impressus (Ameghino, 1887)
Subfamily Hegetotheriinae Ameghino, 1894
Genus Hemihegetotherium Rovereto, 1914
Order Lagomorpha Brandt, 1855
Family Leporidae Fischer, 1817
Genus Lepus Linné, 1758
L. europaeus Linné, 1758
Order Rodentia Bowdich, 1821
Suborder Myomopha Brandt, 1855
Superfamily Muroidea Illiger, 1811
Family Muridae Illiger, 1811
Subfamily Sigmodontinae Wagner, 1843
Tribe Oryzomyini Vorontzov, 1959
Genus Lundomys Voss and Carleton, 1993
Genus Holochilus Brandt, 1835
H. brasiliensis (Desmarest, 1819)
Tribe Akodontini Vorontzov, 1959
Genus Akodon Meyen, 1833
A. azarae (Fischer, 1829)
A. iniscatus Thomas, 1919
Genus Oxymycterus Waterhouse, 1837
Tribe Phyllotini Vorontzov, 1959
Genus Calomys Waterhouse, 1837
C. laucha (Fischer, 1814)
C. musculinus (Thomas, 1913)
Genus Reithrodon Waterhouse, 1837
Reithrodon auritus (Fischer, 1814)
Genus Phyllotis Waterhouse, 1837
Suborder Hysticognathi, Tullberg, 1899
Infraorder Caviomorpha Patterson and Wood in Wood, 1955
Family Octodontidae Waterhouse, 1839
Subfamily Octodontinae Waterhouse, 1839
Genus Pithanotomys Ameghino, 1887
Genus Phtoramys Ameghino, 1887
P. hidalguense Pascual, Pisano and Ortega, 1965
Subfamily Ctenomyinae Tate, 1935
Genus Xenodontomys Kraglievich, 1927
X. ellipticus Kraglievich, 1927
Genus Actenomys Burmeister (in C.V. Burmeister, 1888)
A. priscus (Owen, 1840)
Genus Ctenomys Blainville, 1826
C. kraglievichi (Rusconi, 1930)
C. talarum Thomas, 1898
Family Caviidae Gray, 1821
Subfamily Caviinae Gray, 1821
Genus Palaeocavia Ameghino, 1889
Genus Neocavia Kraglievich, 1932
Genus Dolicavia C. Ameghino, 1916
Genus Galea Meyen, 1833
Genus Microcavia Gervais and Ameghino, 1880
Genus Cavia Pallas, 1766
C. aperea Erxleben, 1777
Subfamily Dolichotinae Pocock, 1922
Genus Orthomyctera Ameghino, 1889
O. lacunosa (Ameghino, 1888)
Family Chinchillidae Bennett, 1833
Genus Lagostomus Brookes, 1828
L. maximus (Desmarest, 1817)
Subgenus Lagostomus (Lagostomopsis) Kraglievich, 1926
Family Myocastoridae Miller and Gidley, 1918
Genus Myocastor Kerr, 1792
M. columnaris Rusconi, 1929
Family Hydrochoeriidae Gill, 1872
Subfamily Hydrochoerinae (Gray, 1825), sensu Kraglievich, 1930
Genus Neochoerus Hay, 1926
N. tarijensis (Ameghino, 1902)
Order Proboscidea Illiger, 1811
Family Gomphotheriidae Hay, 1922
Order Artiodactyla Owen, 1848
Suborder Suiformes Jaeckel, 1911
Infraorder Suina Gray, 1821
Family Tayassuidae Palmer, 1897
Genus Tayassu Fischer, 1814
Suborder Tylopoda Illiger, 1811
Family Camelidae Gray, 1821
Tribe Lamini Webb, 1974
Genus Lama Cuvier, 1800
L. guanicoe (Müller, 1776)
Suborder Ruminantia Scopoli, 1777
Family Cervidae Goldfuss, 1820
Subfamily Odocoileinae Pocock, 1923
Genus Ozotoceros Ameghino, 1891
O. bezoarticus (Linné, 1758)
Family Bovidae Gray, 1821
Subfamily Caprinae Gray, 1821
Genus Ovis Linné, 1758
O. aries Linné, 1758
Subfamily Bovinae Gray, 1821
Genus Bos Linné, 1758
B. taurus Linné, 1758
Family Equidae Gray, 1821
Subfamily Equinae (Gray, 1821) Steinmann and Döderlein, 1890
Genus Equus Linné, 1758
Subgenus E. ( Amerhippus ) Hoffstetter, 195
E. (A.) neogaeus (Lund, 1840)
Genus Hippidion Owen, 1869
H. principale (Lund, 1846)
Order Carnivora Bowdich, 182
Family Canidae Fischer de Waldheim, 1817
Genus Pseudalopex Burmeister, 1856
P. gymnocercus (Fischer, 1814)*
Family Felidae Fischer de Waldheim, 1817
Subfamily Felinae Fischer de Waldheim, 1817
Genus Herpailurus Severtzov, 1858
H. yaguaroundi (Lacépède, 1809)

*Galliari et al . (1996) assign the species of this genus to Lycalopex Burmeister, 1854 following Zunino et al . (1995).

Comments on Appendix 2

The oldest record of Rhea is that of R. anchorenensis , whose type and single specimen (currently lost) was found in the Ensenadan of Punta Anchorena, Buenos Aires Province.
The record of South American iguanians is discontinuous. It includes Cretaceous-Paleocene, Middle Miocene and Pleistocene taxa. They were not recorded so far in the Late Miocene-Pliocene. The fragment of Dique Paso Piedras is the first record for this lapse.
The holotype of Doellotatus inornatus is from Farola Monte Hermoso, without stratigraphic provenance (Rovereto, 1914). Other materials assigned to this species were found in the ?Irenense? of the Quequén Salado river, Buenos Aires province, traditionally assigned to the Chapadmalalan (Goin et al ., 1994) but certain levels of this locality must be considered older than this age (Verzi et al ., 2003). Consequently, its biochron could be older.
Doellotatus praecursor is the most derived species of the genus (Deschamps et al ., 1998). Although this species was recorded in the upper levels of the Río Negro Formation together with Plohophorus aff. figuratus (Aramayo, 1987), the holotype was found at the lower valley of the Río Negro without stratigraphic level (Rovereto, 1914). Other taxa considered typically Huayquerian were found in this area (see Aramayo, 1987). Hence the biochron of D . Praecursor could be older than so far proposed.
Tolypeutes sp. is similar to the specimen found in the Río Salado near Belgrano, Buenos Aires Province, in Bonaerian sediments, that belongs to a new undescribed species ( Tolypeutes n. sp., G. Scillato-Yané pers. com., 2003).
According to Cerdeño and Bond (1998), Paedotherium bonaerense is recorded from the Montehermosan to the Marplatan. The species of the genus Paedotherium are sometimes difficult to determine with isolated fragments because probably they follow an anagenetic evolutionary pattern (M. Bond pers. com., 2003). It may not be discarded that some specimens assigned to P. minor recorded in Epecuén and Arroyo Chasicó formations actually represent small/juvenile specimens of P . bonaerense . Consequently, the biochron of this taxon could have been longer.
The materials of Oxymycterus sp., Lundomys sp. and Phyllotis sp. cited in Alberdi et al . (eds.) 1995, for the Ensenadan are those found in Bajo San José (Pardiñas and Deschamps, 1996), which is considered here Bonaerian. Consequently, their biochron is Bonaerian-Recent.
The materials of Pithanotomys from Chapadmalal (3.3 Ma; Schultz et al ., 1998) are more derived than that of Balneario Saldungaray (D. Verzi pers. com., 2003). Consequently, this latter could be older ( i.e . Montehermosan).
Actenomys is first and abundantly recorded in the Montehermosan in its type locality, and continues to the Chapadmalalan. The systematic of the several known species has not been revisited, but a preliminary revision showed that the specimen of Las Obscuras is similar to those of Farola Monte Hermoso determined as A. priscus (Owen, 1840), and different from those of Chapadmalal (Verzi and Deschamps, unpublished).
Caviids of the Late Miocene and Early Pliocene ( Neocavia , Paleocavia , Dolicavia , Orthomyctera , Lagostomus ( Lagostomopsis )) have to be reviewed, consequently they are not considered for biostratigraphic analyses. Anyway, those specimens of Dolicavia recorded in Grünbein, Cantera Seminario are not as derived as those of the Chapadmalalan (the third lobe of the p4 is smaller and the internal flexid not so deep as in D. minuscula ).
Chinchillids show very low diversity and the only two taxa recognized from the Chasicoan to the Lujanian have not been revisited. Vucetich and Verzi (1995) think there is a single genus for the Plio-Pleistocene chinchillids because differences between Lagostomopsis and Lagostomus are only increasing size, with change in relative proportions of the skull and gradual thinning of cheek teeth. Lagostomus is recorded from the Chasicoan to the Upper Chapadmalalan.Herpailurus yaguaroundi is not cited in the fossil record, but poorly studied materials of Felidae recorded since the Ensenadan are listed as Felis (Cione et al ., 1999).

Appendix 2. (figure 8 ) Range chart of the taxa found in the study area compared to the biochron published in corresponding References. Thin straight line means the known biochron, dotted line means dubious record, and thick straight line means the record in the study area. An asterisk marks those taxa whose biochron is discussed in ?Comments on Appendix 2.? Cs, Chasicoan; Hy, Huayquerian; Mo, Montehermosan; Cp, Chapadmalalan; En, Ensenadan; Bo, Bonaerian; Lu, Lujanian; Pl, Platan; Rc, Recent / Distribución estratigráfica de los taxones hallados en el área de estudio, comparados con los biocrones publicados en las referencias correspondientes. Líneas delgadas: biocrones conocidos; líneas punteadas: registros dudosos; líneas gruesas: registro en el área de estudio. Asterisco: taxones cuyo biocrón se discute en ?Comentarios sobre el Apéndice 2?.

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